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Giventhe need for male and female organs to fit together, theresearcherssuggest that selection of horn size might be the main methodof speciation in Onthophagus. 考慮到雄性和雌性的器官相匹配的規(guī)律，研究人員認為，在食糞甲蟲的角上所發(fā)生的自然選擇作用也許是物種形成的主要途徑。