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Due to the presence of substrate gradients in a bioflim, the amount of OC is difficult to be assessed with experiments. 由于在生物膜內(nèi)存在著基質(zhì)擴(kuò)散梯度，所以，通過試驗(yàn)方法確定凈耗氧量十分困難，甚至難以實(shí)現(xiàn)。